Nine great apes at the San Diego Zoo – four orangutans and five bonobos – made veterinary history in recent weeks as the world’s first non-human primates known to be vaccinated against Covid-19, zoo officials said.

As many as eight gorillas at the San Diego Zoo Safari Park were presumed to have contracted Covid-19 from a human handler after one of the animals tested positive, marking the first known transmission of the virus to apes, zoo officials said.